Gleaning is a feeding strategy by birds and bats in which they catch invertebrate prey, mainly arthropods, by plucking them from foliage or the ground, from crevices such as rock faces and under the eaves of houses, or even, as in the case of ticks and lice, from living animals. This behavior is contrasted with hawking insects from the air or chasing after moving insects such as ants. Gleaning, in birds, does not refer to foraging for seeds or fruit. Gleaning is a common feeding strategy for some groups of birds, including nuthatches, tits (including chickadees), wrens, woodcreepers, treecreepers, Old World flycatchers, Tyrant flycatchers, babblers, Old World warblers, New World warblers, vireos and some hummingbirds and cuckoos. Many birds make use of multiple feeding strategies, depending on the availability of different sources of food and opportunities of the moment.
Techniques and adaptations
Foliage gleaning, the strategy of gleaning over the leaves and branches of trees and shrubs, can involve a variety of styles and maneuvers. Some birds, such as the common chiffchaff of Eurasia and the Wilson's warbler of North America, feed actively and appear energetic. Some will even hover in the air near a leaf or twig while gleaning from it; this behavior is called "hover-gleaning". Other birds are more methodical in their approach to gleaning, even seeming lethargic as they perch upon and deliberately pick over foliage. This behavior is characteristic of the bay-breasted warbler and many vireos. Another tactic is to hang upside-down from the tips of branches to glean the undersides of leaves. Tits such as the familiar black-capped chickadee are often observed feeding in this manner. Some birds, like the ruby-crowned kinglet and red-eyed vireo of North America use a combination of these tactics. Gleaning birds are typically small with compact bodies and have small, sharply pointed bills. These features are even seen in gleaning birds that are not closely related. For example, in flycatchers of the family Tyrannidae, in which some member species are more adapted for hawking insects on the wing and others for gleaning, the gleaners have bills that resemble those of tits and warblers, unlike their larger-billed relatives. Also, some members of the woodpecker family, particularly piculets such as the rufous piculet of Southeast Asia, are similarly adapted for gleaning, with small, compact bodies and sharp bills, rather than the long, supportive tails and wedge-shaped bills more typical of woodpeckers. Birds such as the aforementioned piculet are specialized for gleaning the bark of trees, as are nuthatches, woodcreepers, and treecreepers. Most bark-gleaners work their way up tree trunks or along branches, though nuthatches are well known as birds that can go the opposite direction, facing down and working their way down the trunk, as well. This requires strong legs and feet on the part of the nuthatch and piculet, while birds that face upwards tend to have stiff tail feathers to prop them up. Birds often specialize in a particular niche, such as a particular stratum of forest or type of vegetation. In South and Southeast Asia, for example, the mountain tailorbird is often found gleaning in thickets and stands of bamboo, Abbott's babbler gleans lower-storey foliage in lowland forest, the rufous-chested flycatcher and brown fulvetta are birds of the mid-storey forest, the yellow-breasted warbler gleans in the mid- to upper-storey, and the greater green leafbird specializes in the upper-storey forest. The Javan white-eye is a bird of coastal scrub and mangroves, while the related black-capped white-eye is restricted to montane forest.
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