Gnathovorax is a genus of herrerasaurid saurischian dinosaur from the Santa Maria Formation in Rio Grande do Sul, Brazil. The type and only species is Gnathovorax cabreirai, described by Pacheco et al. in 2019. The type specimen comprises a mostly complete and well-preserved articulated skeleton (considered one of the best herrerasaurid skeletons ever discovered), which fossilized in close association with rhynchosaur and cynodont remains. The discovery of this superb specimen has shed light onto poorly understood aspects of herrerasaurid anatomy such as endocranial soft tissues.
Discovery
The holotype specimen, CAPPA/UFSM 0009, is an almost complete and partially articulated skeleton, lacking only portions of the left shoulder girdle and left forelimb. It was found in 2014 at the Marchezan site, in the municipality of São João do Polêsine, Rio Grande do Sul, Brazil. This locality preserved rocks from Santa Maria Formation in the Candelária Sequence of the Paraná Basin. The skeleton was fossilized within a mudstone layer, along with small articulated skeletons of prozostrodont cynodonts. This mudstone layer was then covered by a layer of sandstone containing disarticulated rhynchosaur remains. These fossil remains are indicative of a diverse community of animals at Marchezan. Stratigraphically correlated beds from a nearby site were dated as the middle of the Carnian stage of the Late Triassic, circa 233.23 ± 0.73 million years ago. The monospecific genus was named and described by Cristian Pacheco, Rodrigo Temp Müller, Max Langer, Flávio Augusto Pretto, Leonardo Kerber and Sérgio Dias da Silva in an article published in 2019. The generic name is derived from the Greek gnathos, jaw, and the Latin vorax, "voracious", from vorō (“devour”) + −āx (“inclined to”). The specific epithet was given in honour of Dr. Sérgio Furtado Cabreira, the palaeontologist who found the specimen.
Description
Skull
There were three teeth in the premaxilla, in contrast to Herrerasaurus (which possessed four). The contact between the premaxilla and maxilla encompassed two fenestrae (holes): a subnarial fenestra which was situated low on the snout and an additional smaller fenestra positioned above it. While a subnarial fenestra is common in herrerasaurids and theropods, the additional fenestra is unique to Gnathovorax. Like other archosaurs, a small basin known as the antorbital fossa is present in front of the antorbital fenestra. Many early saurischians have a hole called the promaxillary fenestra situated on the antorbital fossa, but this hole is absent in Gnathovorax. On the other hand, Gnathovorax does possess a characteristic thin ridge edging an additional emargination within the antorbital fossa; a small depression is present between this ridge and the ridge forming the border of the antorbital fossa. The lower branch of the lacrimal is slender and extends to the middle of the lower edge of the orbit (eye socket), similar to Daemonosaurus but different from Herrerasaurus. Most other aspects of the skull, such as the thick squamosal, expanded front branch of the jugal, the shape of the infratemporal fenestra, and a lack of palatal teeth, are most similar to herrerasaurids among early dinosaurs. The supraoccipital bone at the top of the braincase is trapezoidal when seen from behind (unlike Herrerasaurus's triangular supraoccipital) and has a blade-like crest on top. The occipital condyle is also thicker, the basioccipital has a V-shaped (rather than U-shaped) contact with the parabasisphenoid, and the paroccipital processes are larger in Gnathovorax compared to Herrerasaurus. Otherwise the braincase is similar between the two taxa. The dentary lacks sauropodomorph qualities and instead likely possessed a sliding joint at the chin similar to other herrerasaurids, although preservation is not good enough to fully confirm this. The 3 premaxillary, 19 maxillary, and 14 dentary teeth are all thin, curved, and blade-like. There are serrations on the distal (rear) edge of all the teeth, and the maxillary teeth also have serrations on the mesial (front) edge.
Postcrania
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