Gregor Johann Mendel (; German: [ˈmɛndl̩]; Czech: Řehoř Jan Mendel; 20 July 1822 – 6 January 1884) was an Austrian biologist, meteorologist, mathematician, Augustinian friar and abbot of St. Thomas' Abbey in Brno (Brünn), Margraviate of Moravia. Mendel was born in a German-speaking family in the Silesian part of the Austrian Empire (today's Czech Republic) and gained posthumous recognition as the founder of the modern science of genetics. Though farmers had known for millennia that crossbreeding of animals and plants could favor certain desirable traits, Mendel's pea plant experiments conducted between 1856 and 1863 established many of the rules of heredity, now referred to as the laws of Mendelian inheritance. Mendel worked with seven characteristics of pea plants: plant height, pod shape and colour, seed shape and colour, and flower position and colour. Taking seed colour as an example, Mendel showed that when a true-breeding yellow pea and a true-breeding green pea were cross-bred, their offspring always produced yellow seeds. However, in the next generation, the green peas reappeared at a ratio of 1 green to 3 yellow. To explain this phenomenon, Mendel coined the terms "recessive" and "dominant" in reference to certain traits. In the preceding example, the green trait, which seems to have vanished in the first filial generation, is recessive, and the yellow is dominant. He published his work in 1866, demonstrating the actions of invisible "factors"—now called genes—in predictably determining the traits of an organism. The actual genes were only discovered in a long process that ended in 2025 when the last three of the seven Mendel genes were identified in the pea genome. The profound significance of Mendel's work was not recognized until the turn of the 20th century (more than three decades later) with the rediscovery of his laws. Erich von Tschermak, Hugo de Vries and Carl Correns independently verified several of Mendel's experimental findings in 1900, ushering in the modern age of genetics.
Early life and education Mendel was born into a German-speaking family in Heinzendorf bei Odrau, in Silesia, Austrian Empire (now Hynčice in the Czech Republic). He was the son of Anton and Rosine (Schwirtlich) Mendel and had one older sister, Veronika, and one younger, Theresia. They lived and worked on a farm which had been owned by the Mendel family for at least 130 years (the house where Mendel was born is now a museum devoted to Mendel). During his childhood, Mendel worked as a gardener and studied beekeeping. As a young man, he attended gymnasium in Troppau (Czech: Opava). Due to illness, he had to take four months off during his gymnasium studies. From 1840 to 1843, he studied practical and theoretical philosophy and physics at the Philosophical Institute of the University of Olomouc (German: Olmütz), taking another year off because of illness. He also struggled financially to pay for his studies, and Theresia gave him her dowry. Later, he helped support her three sons, two of whom became doctors. He became a monk partly because it enabled him to obtain an education without paying for it himself. As the son of a struggling farmer, the monastic life, in his words, spared him the "perpetual anxiety about a means of livelihood." Born Johann Mendel, he was given the name "Gregor" (Řehoř in Czech) when he joined the Order of Saint Augustine.
Academic career
When Mendel entered the Faculty of Philosophy, the Department of Natural History and Agriculture was headed by Johann Karl Nestler, who conducted extensive research on hereditary traits of plants and animals, especially sheep. Upon recommendation of his physics teacher Friedrich Franz, Mendel entered the Augustinian St Thomas's Abbey in Brno and began his training as a Catholic priest. Mendel worked as a substitute high school teacher. In 1850, he failed his exams' oral part, the last of three parts, to become a certified high school teacher. In 1851, he was sent to the University of Vienna to study under the sponsorship of Abbot Cyril František Napp so that he could get a more formal education. At Vienna, his professor of physics was Christian Doppler. Mendel returned to his abbey in 1853 as a teacher, principally of physics. In 1854, he met Aleksander Zawadzki, who encouraged his research in Brno. In 1856, he took the exam to become a certified teacher and again failed the oral part. In the summer of 1862, he joined an organised group tour to Paris and London, where he visited the International Exhibition and major scientific sites, a trip that may have influenced the final stage of his hybridisation research. In 1867, he succeeded Napp as abbot of the monastery. After he was elevated as abbot in 1868, his scientific work largely ended, as Mendel became overburdened with administrative responsibilities, especially a dispute with the civil government over its attempt to impose special taxes on religious institutions. Mendel died on 6 January 1884, at the age of 61, in Brno, from chronic nephritis. Czech composer Leoš Janáček played the organ at his funeral. After his death, the succeeding abbot burned all papers in Mendel's collection, to mark an end to the disputes over taxation. The exhumation of Mendel's corpse in 2021 delivered some physiognomic details like body height (168 cm (66 in)). His genome was analysed, revealing that Mendel was predisposed to heart problems.
Contributions
Experiments on plant hybridization
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