Mediterranean forests, woodlands and scrub is a biome defined by the World Wide Fund for Nature. The biome is generally characterized by dry summers and rainy winters, although in some areas rainfall may be uniform. Summers are typically hot in low-lying inland locations, but can be cool near colder seas; while winters are typically mild to cool in low-lying locations, but can be cold in inland and higher locations. These ecoregions are highly distinctive, collectively harboring 10% of the Earth's plant species.
Distribution
The Mediterranean forests, woodlands, and scrub biome mostly occurs in, but is not limited to, the Mediterranean climate zones of the mid-latitudes:
the Mediterranean Basin the Chilean Matorral the California chaparral and woodlands the Fynbos of South Africa the Mallee Woodlands and Shrublands of Australia The biome is not limited to the Mediterranean climate zone. It can also be present in other climate zones (which typically border the Mediterranean climate zone), such as the drier regions of the oceanic and humid subtropical climates, and as well as the lusher areas of the semi-arid climate zone. Non-Mediterranean climate regions that would feature Mediterranean vegetation include the Nile River Valley in Egypt (extending upstream along the riverbanks), parts of the Eastern Cape in South Africa, southeastern Australia, southeastern Azerbaijan, southeastern Turkey, far northern Iraq, the Mazandaran Province in Iran, Central Italy, parts of the Balkans (including Northern Greece), Southern Crimea in Ukraine, as well as Northern and Western Jordan.
Vegetation
Vegetation types range from forests to woodlands, savannas, shrublands, and grasslands; "mosaic habitat" landscapes are common, where differing vegetation types are interleaved with one another in complex patterns created by variations in soil, topography, exposure to wind and sun, and fire history. Much of the woody vegetation in Mediterranean-climate regions is sclerophyll, which means 'hard-leaved' in Greek. Sclerophyllous vegetation generally has small, dark leaves covered with a waxy outer layer to retain moisture in the dry summer months. Phytogeographers consider the fynbos (South Africa) as a separate floral kingdom because 68% of the 8,600 vascular plant species crowded into its 90,000 square kilometers (35,000 mi2) are endemic and highly distinctive at several taxonomic levels. This is equivalent to about 40% of the plant species of the United States and Canada combined, found within an area the size of the state of Maine. The fynbos and Southwest Australia shrublands have flora that are significantly more diverse than the other ecoregions, although any Mediterranean shrubland is still rich in species and endemics relative to other non-forest ecoregions.
Biome plant groups
Major plant communities in this biome include:
Forest: Mediterranean forests are generally composed of broadleaf trees, such as the oak and mixed sclerophyll forests of California and the Mediterranean region, the Eucalyptus forests of Southwest Australia, and the Nothofagus forests of central Chile. Forests are often found in riparian areas, where they receive more summer water. Coniferous forests also occur, especially around the Mediterranean. Pine and deciduous oak forest are widespread across California. Pinus halepensis, lentisk (Pistacia lentiscus), kermes oak (Quercus coccifera) and Chamaerops are found across Spanish Mediterranean forest. Woodland: Oak woodlands are characteristic of the Mediterranean Basin and in California. Pine woodlands are also present in the Mediterranean Basin. California additionally has walnut woodlands. Savanna and grassland: The California Central Valley grasslands are the largest Mediterranean grassland eco-region, although these grasslands have mostly been converted to agriculture. The remaining woodlands feature mainly oak, walnut and pine. The cork oak savanna in Portugal, known as montado, is a good example of a mediterranean savanna. Shrubland: Shrublands are dense thickets of evergreen sclerophyll shrubs and small trees. They are most common near the seacoast, and are often adapted to wind and salt air from the ocean. They are called chaparral (California and southern Portugal), matorral in Chile and southern Spain, garrigue or maquis in France, macchia or gariga in Italy, phrygana in Greece, tomillares in Spain, fynbos, renosterveld, Succulent Karoo, and strandveld in South Africa, kwongan in Southwest Australia and batha in Israel. Northern coastal scrub and coastal sage scrub, also known as soft chaparral, occur near the California coast. In some places shrublands are of the mature vegetation type, and in other places are the result of degradation of former forest or woodland by logging or overgrazing, or disturbance by major fires.
Fire as a medium of change
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