Nanohalarchaeota is a phylum of diminutive archaea with small genomes and limited metabolic capabilities, belonging to kingdom Nanobdellati. They are ubiquitous in hypersaline habitats, which they share with the extremely halophilic haloarchaea. Nanohaloarchaea were first identified from metagenomic data as a class of uncultivated halophilic archaea composed of six clades and were subsequently placed in the phylum Nanohalarchaeota within the DPANN clade (kingdom Nanobdellati). However, the phylogenetic position of nanohaloarchaea is still highly debated, being alternatively proposed as the sister-lineage of haloarchaea or a member of the Nanobdellati kingdom. The lineage has since been identified in data from a range of hypersaline environments including: Australian thalassohaline lake, Spanish saltern, Russian soda brine, Californian saltern, Chilean halite, and Ethiopian Dallol hydrothermal system. Like most other members of DPANN, they are only known to live with a host (specifically a similarly halophilic archaeon) in symbiosis. The exact nature of this interaction with the host vary among species, from mutualism to parasitic predation.
Viruses Several species of nanhoaloarchaea encode CRISPR-Cas systems suggesting that they are parasitized by viruses. Consistently, metagenomic analysis has revealed that nanohaloarchaea are associated with diverse viruses. These viruses belong to four morphologically different groups, including those with head-tailed (class Caudoviricetes), tailless icoahedral (realm Singelaviria), enveloped pleomorphic and spindle-shaped virions. Currently known nanohaloarchaeal viruses have been classified into seven distinct families.
Taxonomy The currently accepted taxonomy is based on the List of Prokaryotic names with Standing in Nomenclature (LPSN) and National Center for Biotechnology Information (NCBI).
Phylum "Nanohalarchaeota" corrig. Rinke et al. 2013
Class "Nanohalarchaeia" corrig. Narasingarao et al. 2012 Order "Nanohalarchaeales" Family "Nanohalarchaeaceae" Genus ?"Candidatus Nanohalarchaeum" corrig. Hamm et al. 2019 "Ca. N. antarcticum" corrig. Hamm et al. 2019 Genus ?"Candidatus Nanopetraeus" corrig. Crits‐Christoph et al. 2016 ["Nanopetramus" (sic)] "Ca. N. primus" corrig. Crits‐Christoph et al. 2016 Order "Nanohydrothermales" Xie et al. 2022 Family "Nanohydrothermaceae" Xie et al. 2022 Genus ?"Candidatus Nanohydrothermus" Xie et al. 2022 "Ca. N. guaymasensis" Xie et al. 2022 Order "Nucleotidisoterales" Xie et al. 2022 Family "Nanosalenecaceae" Xie et al. 2022 Genus ?"Candidatus Nanosalenecus" Xie et al. 2022 "Ca. N. halilacustris" Xie et al. 2022 Family "Nucleotidisoteraceae" Xie et al. 2022 Genus ?"Candidatus Nucleotidisoter" Xie et al. 2022 "Ca. N. xinjiangensis" Xie et al. 2022 Family "Nucleotidivindicaceae" Xie et al. 2022 Genus ?"Candidatus Nucleotidivindex" Xie et al. 2022 "Ca. N. qijiaojingensis" Xie et al. 2022 Class "Nanohalobiia" corrig. La Cono et al. 2020 ["Nanosalinia" Rinke et al. 2021] Order "Nanohalobiales" La Cono et al. 2020 ["Nanosalinales" Rinke et al. 2020] Family "Nanoanaerosalinaceae" Zhao et al. 2022 Genus "Candidatus Asbonarchaeum" Baker et al. 2024 "Ca. A. danakilense" Baker et al. 2024 Genus "Candidatus Nanoanaerosalina" Zhao et al. 2022 "Ca. N. halalkaliphila" Zhao et al. 2022 Family "Nanohalalkaliarchaeaceae" Zhao et al. 2022 Genus "Candidatus Nanohalalkaliarchaeum" Zhao et al. 2022 "Ca. N. halalkaliphilum" Zhao et al. 2022 Family "Nanohalobiaceae" La Cono et al. 2020 ["Nanosalinaceae" Rinke et al. 2020] Genus ?"Candidatus Haloredivivus" Ghai et al. 2011 "Ca. N. primus" corrig. Ghai et al. 2011 Genus "Candidatus Nanohalobium" La Cono et al. 2020 "Ca. N. constans" La Cono et al. 2020 Genus "Candidatus Nanohalococcus" Reva et al. 2023 "Ca. N. occultus" Reva et al. 2023 Genus "Candidatus Nanohalovita" Reva et al. 2023 "Ca. N. haloferacivicina" Reva et al. 2023 Genus "Candidatus Nanosalina" Narasingarao et al. 2012 "Ca. N. primus" corrig. Narasingarao et al. 2012 Genus "Candidatus Nanosalinicola" corrig. Narasingarao et al. 2012 ["Nanosalinarum" (sic)] "Ca. N. primus" corrig. Narasingarao et al. 2012
See also List of Archaea genera
References
Further reading Ghai; et al. (2011). "New Abundant Microbial Groups in Aquatic Hypersaline Environments". Scientific Reports. 1 (135) 135. Bibcode:2011NatSR...1..135G. doi:10.1038/srep00135. PMC 3216616. PMID 22355652. Narasingarao; et al. (2012). "De novo metagenomic assembly reveals abundant novel major lineage of Archaea in hypersaline microbial communities". ISME J. 6 (1): 81–93. Bibcode:2012ISMEJ...6...81N. doi:10.1038/ismej.2011.78. PMC 3246234. PMID 21716304.
