Neottia is a genus of orchids. The genus now includes the former genus Listera, commonly known as twayblades, referring to the single pair of opposite leaves at the base of the flowering stem. The genus is native to temperate, subarctic and arctic regions across most of Europe, northern Asia (Siberia, China, the Himalayas, Central Asia, etc), and North America, with a few species extending into subtropical regions in the Mediterranean, Indochina, the southeastern United States, etc. Neottia produces a racemose inflorescences with flowers in shades of green or dull pink through to maroon and purple. The lip of each flower is prominently forked or two-lobed. Some species (those which were previously the only members of the genus Neottia in the strict sense, such as the bird's-nest orchid, Neottia nidus-avis) are completely without chlorophyll and have leaves which are reduced to scales.
Description Neottia is a genus of relatively small terrestrial orchids. Some (the former genus Listera) have chlorophyll and are hence gaining their energy from photosynthesis. Others (the formerly narrowly defined genus Neottia) lack chlorophyll and are dependent on fungi for their nutrition (mycotrophic). The flowering stem has a number of greenish or brownish bracts at the base. In the photosynthetic members of the genus there are also two more-or-less opposite green leaves (very rarely more than two in Neottia ovata). The flowers are individually small, in shades of green, yellow, brown or red to purple. The lip is usually much larger than the other five tepals, and is almost always deeply divided into two lobes at the end. The other five tepals may form a loose hood. The pollinia are not stalked.
Taxonomy At one time the genus was divided between Neottia and Listera. Molecular phylogenetic studies in this century have shown that species lacking chlorophyll, such as Neottia nidus-avis, evolved within a larger clade of photosynthetic plants containing Neottia and Listera, so that the two genera should be combined. As Neottia is the older name, sources such as the World Checklist of Selected Plant Families and the Flora of China now use Neottia for all species formerly in Listera. Other sources continued to divide the genus into two.
Species As of May 2026, Plants of the World Online accepted the following species:
Neottia acuminata Schltr. Neottia alternifolia (King & Pantl.) Szlach. Neottia anthropophora (Aver. & V.C.Nguyen) R.Kr.Singh & Sanjeet Kumar Neottia atayalica T.C.Hsu Neottia auriculata (Wiegand) Szlach. Neottia bambusetorum (Hand.-Mazz.) Szlach. Neottia banksiana (Lindl.) Rchb.f. Neottia bicallosa X.H.Jin Neottia biflora (Schltr.) Szlach. Neottia bifolia (Raf.) Baumbach Neottia borealis (Morong) Szlach. Neottia brevicaulis (King & Pantl.) Szlach. Neottia brevilabris Tang & F.T.Wang Neottia breviscapa T.C.Hsu Neottia camtschatea (L.) Spreng. Neottia chandrae Raskoti, J.J.Wood & Ale Neottia chawalongensis J.D.Ya & D.Z.Li Neottia chenii S.W.Gale & P.J.Cribb Neottia cinsbuensis T.P.Lin & D.M.Huang Neottia confusa Bhaumik Neottia convallarioides (Sw.) Rich. Neottia cordata (L.) Rich. Neottia deltoidea (Fukuy.) Szlach. Neottia dentata (King & Pantl.) Szlach. Neottia dihangensis Bhaumik Neottia divaricata (Panigrahi & P.Taylor) Szlach. Neottia fangii (Tang & F.T.Wang ex S.C.Chen & G.H.Zhu) S.C.Chen, S.W.Gale & P.J.Cribb Neottia flabellata (W.W.Sm.) Szlach. Neottia fugongensis (X.H.Jin) J.M.H.Shaw Neottia fukuyamae T.C.Hsu & S.W.Chung Neottia furusei T.Yukawa & Yagame Neottia gaudissartii Hand.-Mazz. Neottia hohuanshanensis T.P.Lin & Shu H.Wu Neottia inagakii Yagame, Katsuy. & T.Yukawa Neottia inayatii (Duthie) Schltr. Neottia japonica (Blume) Szlach. Neottia karoana Szlach. Neottia kiusiana T.Hashim. & S.Hatus. Neottia kuanshanensis H.J.Su Neottia latilabra (Evrard ex Gagnep.) M.H.J.van der Meer Neottia lihengiae J.D.Ya, H.Jiang & D.Z.Li Neottia linzhiensis X.H.Jin Neottia listeroides Lindl. Neottia longicaulis (King & Pantl.) Szlach. Neottia mackinnonii Deva & H.B.Naithani Neottia makinoana (Ohwi) Szlach. Neottia maolanensis M.N.Wang Neottia megalochila S.C.Chen Neottia meifongensis (H.J.Su & C.Y.Hu) T.C.Hsu & S.W.Chung Neottia microauriculata T.C.Hsu Neottia microglottis (Duthie) Schltr. Neottia microphylla (S.C.Chen & Y.B.Luo) S.C.Chen, S.W.Gale & P.J.Cribb Neottia milinensis J.J.Lei & X.H.Jin Neottia morrisonicola (Hayata) Szlach. Neottia motuoensis X.H.Jin Neottia mucronata (Panigrahi & J.J.Wood) Szlach. Neottia nanchuanica (S.C.Chen) Szlach. Neottia nandadeviensis (Hajra) Szlach. Neottia nankomontana (Fukuy.) Szlach. Neottia nepalensis (N.P.Balakr.) Szlach. Neottia nidus-avis (L.) Rich. Neottia nipponica (Makino) Szlach. Neottia nujiangensis X.H.Jin Neottia nyinyikyawii X.H.Jin, A.T.Mu & H.A.Mung Neottia oblata (S.C.Chen) Szlach. Neottia ovata (L.) Hartm. Neottia pantlingii (W.W.Sm.) Tang & F.T.Wang Neottia papilligera Schltr. Neottia pekinensis (X.Y.Mu & Bing Liu) R.Kr.Singh & Sanjeet Kumar Neottia piluchiensis T.P.Lin Neottia pinetorum (Lindl.) Szlach. Neottia pseudonipponica (Fukuy.) Szlach. Neottia puberula (Maxim.) Szlach. Neottia ruisuiensis T.P.Lin Neottia shenlengiana T.C.Hsu Neottia smallii (Wiegand) Szlach. Neottia smithiana Schltr. Neottia smithii (Schltr.) Szlach. Neottia taibaishanensis P.H.Yang & K.Y.Lang Neottia taizanensis (Fukuy.) Szlach. Neottia tatakaensis T.C.Hsu & H.C.Hung Neottia tenii Schltr. Neottia tenuis (Lindl.) Szlach. Neottia unguiculata (W.W.Sm.) Szlach. Neottia × veltmanii (Case) Baumbach Neottia wardii (Rolfe) Szlach. Neottia wuyishanensis B.Hua Chen & X.H.Jin Neottia yunnanensis (S.C.Chen) Szlach.
References
External links
Jepson Manual Treatment (former Listera only)



