Cathartidae, commonly known as New World vultures and condors, are a family of birds of prey consisting of seven extant species in five genera. It includes five extant vultures and two extant condors found in the Americas. They are known as "New World" vultures to distinguish them from Old World vultures, with which the Cathartidae does not form a single clade despite the two being similar in appearance and behavior as a result of convergent evolution. Like other vultures, New World vultures are scavengers, having evolved to feed off of the carcasses of dead animals without any notable ill effects. Some species of New World vulture (Cathartes sp.) have a good sense of smell, whereas Old World vultures find carcasses exclusively by sight. Other adaptations shared by both Old and New World vultures include a bald head, devoid of feathers which helps prevent rotting matter from accumulating while feeding, and an extremely disease-resistant digestive system to protect against dangerous pathogens found in decaying meat.
Taxonomy and systematics
The family Cathartidae was introduced (as the subfamily Cathartinae) by the French ornithologist Frédéric de Lafresnaye in 1839. The New World vultures comprise seven species in five genera, being Coragyps, Cathartes, Gymnogyps, Sarcoramphus, and Vultur. Of these, only Cathartes is not monotypic. The family's scientific name, Cathartidae, comes from cathartes, Greek for "purifier". Although New World vultures and Old World vultures are not very closely related, they share many resemblances because of convergent evolution. Phylogenetic analyses including all Cathartidae species found two primary clades. The first consists of black vultures (Coragyps atratus) together with the three Cathartes species (lesser yellow-headed vultures (C. burrovianus), greater yellow-headed vultures (C. melambrotus), and turkey vultures (C. aura)), while the second consists of king vultures (Sarcoramphus papa), California condors (Gymnogyps californianus) and Andean condors (Vultur gryphus). New World vultures were traditionally placed in a family of their own in the Falconiformes. However, in the late 20th century some ornithologists argued that they are more closely related to storks on the basis of karyotype. Thus some authorities placed them in the Ciconiiformes with storks and herons; Sibley and Monroe (1990) even considered them a subfamily of the storks. This was criticized and an early DNA sequence study was based on erroneous data and subsequently retracted. However, a 2021 analysis of mitochondrial genes suggested a stronger phylogenetic relationship between Cathartiformes and Accipitriformes, New World vultures were previously placed in the order Cathartiformes. However, a 2021 analysis of mitochondrial genes suggested a stronger phylogenetic relationship between Cathartiformes and Accipitriformes and they are now placed in the same order as Old World vultures. Recent multi-locus DNA studies on the evolutionary relationships between bird groups indicate that New World vultures are related to the other birds of prey, excluding the Falconidae. This analysis argues that New World vultures should either be a part of the new order Accipitriformes or part of the order Cathartiformes closely related to, but distinct from, other birds of prey. Many taxonomists, including the IOC, now agree that New World vultures are part of Accipitriformes, along with families Accipitridae (buzzards, eagles, harriers, hawks, kites, and Old World vultures), Pandionidae (osprey), and Sagittariidae (secretarybird). However, the American Ornithological Society, Handbook of the Birds of the World (HBW) taxonomy, and eBird/Clements taxonomy maintain Cathartidae as the only family within order Cathartiformes. These families are basal members of the recently recognized clade Afroaves.
Extinct species and fossils The fossil history of the Cathartidae is complex, and many taxa that may possibly have been New World vultures have at some stage been treated as early representatives of the family. There is no unequivocal European record from the Neogene.
It is clear that the Cathartidae had a much higher diversity in the Plio-Pleistocene, rivalling the current diversity of Old World vultures and their relatives in shapes, sizes, and ecological niches. Extinct taxa include:
Gobicathartes Late Eocene of Mongolia Diatropornis Late Eocene/Early Oligocene – ?Middle Oligocene of France Phasmagyps Chadronian of Colorado Brasilogyps Late Oligocene/Early Miocene of Brazil Hadrogyps Middle Miocene of SW North America Cathartidae gen. et sp. indet. Late Miocene/Early Pliocene of Lee Creek Mine, US Pliogyps Late Miocene – Late Pliocene of S North America Perugyps Pisco Late Miocene/Early Pliocene of SC Peru Dryornis Early – Late? Pliocene of Argentina; may belong to modern genus Vultur Cathartidae gen. et sp. indet. Middle Pliocene of Argentina Aizenogyps Late Pliocene of SE North America Breagyps Late Pleistocene of SW North America Geronogyps Late Pleistocene of Argentina and Peru Gymnogyps varonai Late Quaternary of Cuba Wingegyps Late Pleistocene of Brazil Pleistovultur Late Pleistocene/Early Holocene of Brazil Cathartidae gen. et sp. indet. Cuba Gymnogyps amplus Late Pleistocene – Holocene of W North America Kuntur cardenasi (Stucchi et al. 2015)
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