The North American beaver (Castor canadensis) is one of two extant beaver species, along with the Eurasian beaver (Castor fiber). It is native to North America and has been introduced in South America (Patagonia) and Europe (primarily Finland and Karelia). The North American beaver is one of the national symbols of Canada and the official state mammal of Oregon and New York. North American beavers are widespread across the continental United States, Canada, southern Alaska, and some parts of northern Mexico. In Canada and the United States, the North American beaver is often referred to simply as "beaver", although this can cause some confusion because another distantly related rodent, Aplodontia rufa, is often called the "mountain beaver". Other vernacular names, including American beaver and Canadian beaver, distinguish this species from the other extant beaver species, Castor fiber, which is native to Eurasia.
Taxonomy
Evolution The first fossil records of beavers are 10 to 12 million years old in Germany, and they are thought to have migrated to North America across the Bering land bridge. During the mid to late Pleistocene epoch, giant beavers weighing up to 200 pounds thrived in central North America. These large early beavers were not yet known to fell trees and build dams. The oldest fossil record (Castor californicus) of beavers in North America are of two beaver teeth near Dayville, Oregon, and are 7 million years old. Early beavers had diverse ways of life, taking on terrestrial and semi aquatic forms. Dam building originated over 10 million years ago by a semi-aquatic variation of beavers.
Subspecies At one time, 25 subspecies of beavers were identified in North America, with distinctions based primarily on slight morphological differences and geographical isolation at the time of discovery. However, modern techniques generally use genetics rather than morphology to distinguish between subspecies, and currently the Integrated Taxonomic Information System (which provides authoritative taxonomic information on plants, animals, fungi, and microbes of North America and the world) does not recognize any subspecies of C. canadensis, though a definitive genetic analysis has not been performed. Such an analysis would be complicated by the fact that substantial genetic mixing of populations has occurred because of the numerous reintroduction efforts intended to help the species recover following extirpation from many regions. The most widespread (formerly recognized) subspecies, which perhaps are now best thought of as populations with some distinct physical characteristics, are C. c. acadicus (New England beaver), C. c. canadensis (Canadian beaver), C. c. carolinensis (Carolina beaver), and C. c. missouriensis (Missouri River beaver). The Canadian beaver originally inhabited almost all of the forested area of Canada, and because of its more valued fur, was often selected for reintroductions elsewhere. The Carolina beaver is found in the southeastern United States; the Missouri River beaver, as its name suggests, is found in the Missouri River and its tributaries; and C. c. acadicus is found throughout the New England area in the northeastern United States.
Description
The beaver is the largest rodent in North America and competes with its Eurasian counterpart, the European beaver, for being the third-largest in the world, both following the South American capybara and lesser capybara. The European species is slightly larger on average but the American has a larger known maximum size. Adults usually weigh from 11 to 32 kg (24 to 71 lb), with 20 kg (44 lb) being typical. In New York, the average weight of adult male beavers was 18.9 kg (42 lb), while non-native females in Finland averaged 18.1 kg (40 lb). However, adults of both sexes averaged 16.8 kg (37 lb) in Ohio. The species seems to conform to Bergmann's rule, as northern animals appear to be larger. In the Northwest Territory, adults weighed a median of 20.5 kg (45 lb). The American beaver is slightly smaller in average body mass than the Eurasian species. The head-and-body length of adult North American beavers is 74–90 cm (29–35 in), with the tail adding a further 20–35 cm (7.9–13.8 in). Very old individuals can exceptionally exceed normal sizes, weighing more than 40 kg (88 lb) or even as much as 50 kg (110 lb). The beavers are semiaquatic with physical traits suited to this lifestyle. It has a large, scaly, paddle-shaped tail and webbed hind feet. The unwebbed front feet are smaller, have claws, and are highly dexterous. Beavers use their front feet for digging, to feed themselves, and to maneuver small stems as they gnaw bark off trees. The eyes are covered by a nictitating membrane which allows the beaver to see underwater. The nostrils and ears are sealed while submerged. Their lips can be closed behind their front teeth so that they can continue to gnaw underwater. A thick layer of fat under its skin provides insulation. The beaver's fur consists of long, coarse outer hairs and short, fine inner hairs, and is usually dark brown. Scent glands near the genitals secrete an oily substance known as castoreum, which the beaver uses to waterproof its fur. There is also another set of oil glands producing unique chemical identifiers in the form of waxy esters and fatty acids. The beaver, like all rodents, possesses continuously growing incisors, and is a hindgut fermenter whose cecum, populated by symbiotic bacteria, helps to digest plant-based material. The beaver is specialized for efficient digestion of its lignocellulose-heavy diet.
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