Papilio machaon, the Old World swallowtail, is a butterfly of the family Papilionidae. The butterfly is also known as the common yellow swallowtail or simply the swallowtail (a common name applied to all members of the family, but this species was the first to be given the name). It is the type species of the genus Papilio. This widespread species is found in much of the Palearctic and in North America.
Etymology This species is named after Machaon (Ancient Greek: Μαχάων, romanized: Makháōn) a figure in Greek mythology. He was a son of Asclepius in the works of Homer.
Taxonomy Papilio machaon was named by Carl Linnaeus in the 10th edition of Systema Naturae in 1758, alongside nearly 200 other species of butterfly. Later, Pierre André Latreille designated it as the type species of the genus Papilio.
Subspecies There are 41 recognized subspecies, that include:
Papilio machaon gorganus is strongly migratory in Europe and may be found in almost all habitats. In the UK, P. m. britannicus is an endemic subspecies, but occasionally individuals of the continental subspecies P. m. gorganus breed temporarily on the south coast. Subspecies P. m. britannicus differs from the continental subspecies in being more heavily marked in black. The Maltese Islands are home to another endemic subspecies, P. m. melitensis.
Distribution and status
This butterfly is present throughout the entire Palearctic region, ranging from Russia to China and Japan, (including the Himalayas and Taiwan), and across into Alaska, Canada, and the United States, and thus, is not restricted to the Old World, despite the common name. In Asia, it is reported as far south as Saudi Arabia, Oman, the high mountains of Yemen, Lebanon, Iran and Israel. In southern Asia, it occurs in Pakistan and Kashmir, northern India (Sikkim, to Assam, and Arunachal Pradesh), Nepal, Bhutan, and northern Myanmar. This butterfly is widespread in Europe. In the United Kingdom, it is limited to a few areas in the Norfolk Broads of East Anglia. It is the UK's largest resident butterfly. The monarch (Danaus plexippus) is slightly larger, but is only a rare vagrant. As P. machaon is widespread throughout Eurasia and often common, it is not threatened as a species. It is listed as "vulnerable" in the South Korean and Austrian Red Data Books, and in the Red Data Book of the former Soviet Union. In Armenia the species demonstrates stable population trend and is assessed as Least Concern. In some countries, P. machaon and its subspecies are protected by law. Papilio machaon machaon is protected by law in six provinces of Austria, Czech Republic, Slovakia, Hungary, Romania, and Moldova. The species is protected in the United Kingdom, and subspecies verityi is protected in India. In late 2017, P. machaon was among several species of butterflies selected by the Estonian Society of Lepidopterists as contenders for the National Butterfly of Estonia. Nearly 5,000 members of the public voted online, with P. machaon receiving 2,664 votes, overwhelmingly winning the title. As well as becoming the National Butterfly of Estonia, P. machaon was named as the Butterfly of the Year for 2018; an honorary title given to a native butterfly species in Estonia annually.
Description The imago typically has yellow wings with black vein markings, and a wingspan of 65–86 millimetres (2.6–3.4 in). The hindwings of both sexes have a pair of protruding tails which give the butterfly its common name from the resemblance to the birds of the same name. Just below each tail is one red and six blue eye spots. In the caterpillar stage, P. machaon has a length of 45 millimetres (1.8 in). When young, the caterpillar resembles a bird dropping, giving it camouflage. The caterpillar also protects itself using a large orange fork which protrudes behind its head. It can be distinguished from Papilio hospiton, which occurs sympatrically with it on Corsica and Sardinia, by the longer "tails" on the hindwings. It can be told apart from the Algerian species Papilio saharae only by counting the segments on the antennae.
Ecology
The butterfly has a strong and fast flight, but frequently pauses to hover over flowering herbs and sip nectar. It frequents alpine meadows and hillsides, and males are fond of 'hilltopping', congregating near summits to compete for passing females. At lower elevations, it can be seen visiting gardens. In Central Europe, the species frequently colonizes cultural landscapes and domestic gardens where host plants are available. Female butterflies are often observed ovipositing on young sprouts of cultivated umbellifers such as dill (Anethum graveolens) or parsley (Petroselinum crispum), even in early spring (April). These plants provide a high-quality nutritional source for the first generation of caterpillars. Unlike other swallowtails which specialise on Rutaceae, this species mostly feeds on plants of family Umbelliferae, females laying eggs singly. Milk parsley (also known as marsh hog's fennel) is normally the only food plant used by the caterpillars of the British subspecies. The food plants of the swallowtail in Europe, Asia, and North America are more varied than in the UK. It uses a wide variety of umbellifers including wild carrot (Daucus carota), wild angelica (Angelica sylvestris), fennel (Foeniculum vulgare), and hogweeds (Heracleum). In the Maltese Islands, the caterpillar feeds on plants such as rue (Ruta chalepensis) in addition to Umbelliferae such as fennel. In Kashmir, the common yellow swallowtail, as Papilio machaon is called there, inhabits alpine meadows in the Himalayas occurring from 2,000 feet (610 m) in Kashmir valley to 16,000 ft (4,900 m) in the Garhwal Himalayas. In India, in Himachal Pradesh, it is found over 4,000 ft (1,200 m) only and in Sikkim over 8,000 ft (2,400 m) only. At lower elevations, these butterflies fly from March to September; at higher elevations, they are limited by the short summer seasons. The British subspecies P. m. brittanicus is less mobile than its European continental counterpart and stays within, or close by, its fenland habitat.
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