The Philippine tarsier (Carlito syrichta) is a species of tarsier endemic to the Philippines. It is found in the southeastern part of the archipelago, particularly on the islands of Bohol, Samar and Leyte. It is a member of the approximately 45-million-year-old family Tarsiidae, whose name is derived from its elongated "tarsus" or ankle bone. Formerly a member of the genus Tarsius, it has since been listed as the only member of the genus Carlito, a new genus named after the conservationist Carlito Pizarras. Its geographic range also includes Maripipi Island, Siargao Island, Basilan Island and Dinagat Island. Tarsiers have also been reported in Sarangani, although they may be different subspecies.
Etymology and taxonomic history
The tarsier is named for its elongated "tarsus" or ankle bone. This tarsier is known locally as mawumag in Visayan languages including Cebuano, and magô in Waray. It is also known as mamag, magau, malmag, and magatilok-iok.
Taxonomic classification The Philippine tarsier is the only member of the genus Carlito and a member of the family Tarsiidae. Three subspecies are recognised:
Family Tarsiidae Genus Tarsius: found in Sulawesi Genus Cephalopachus: found in Sundaland Genus Carlito: found in Greater Mindanao Species Carlito syrichta Subspecies Carlito syrichta syrichta from Leyte and Samar Subspecies Carlito syrichta fraterculus from Bohol Subspecies Carlito syrichta carbonarius from Mindanao
Changes in taxonomy Previously all living tarsiers had been placed in the genus Tarsius, but a 2010 taxonomic revision by Shekelle and Groves placed the distinctive Philippine tarsier in its own genus, Carlito named after Filipino conservationist Carlito Pizarras. The Philippine tarsier was introduced to Western biologists in the 18th century through the missionary J.G. Camel's description given to J. Petiver of an animal said to have come from the Philippines. Petiver published Camel's description in 1705 and named the animal Cercopithecus luzonis minimus which was the basis for Linnaeus' (1758) Simia syrichta and eventually Carlito syrichta, the scientific name as of 2010. The IUCN taxonomic note lists two subspecies, but the non-nominate one is poorly defined at present, so the species is treated as a whole. Tarsius syrichta carbonarius and Tarsius s. fraterculus were recognized by Hill (1955) as weakly defined subspecies. Niemitz (1984) found the differences to be insignificant based upon comparisons with museum specimens. Musser and Dagosto (1987) felt that the available museum specimens were insufficient to resolve the issue, but mentioned that Heaney felt that a single male tarsier from Dinagat might be distinct. Groves (2001) did not recognize any subspecies of C. syrichta, but Groves and Shekelle (2010) recognized the subspecies C. s. fraterculus, C. s. syrichta, and C. s. carbonarius when splitting the species of Tarsius into Carlito.
Anatomy and morphology The Philippine tarsier measures 85 to 160 mm (3.35 to 6.30 in) in height, making it one of the smallest primates. The small size makes it difficult to spot. The mass for males is between 80–160 g (2.8–5.6 oz), usually lighter for females, somewhat heavier than other tarsiers such as the pygmy tarsier. The average adult is about the size of an adult human fist. The female tarsier has multiple sets of breasts, but the only functional set is at the pectoralis. The other breasts are used as anchor points for the newborn tarsiers. The gestation period lasts 180 days, or 6 months, after which only one tarsier is born. The newborn tarsier is born with much fur and eyes open. Its body and head length is about 70 mm (2.8 in), and its tail is around 115 mm (4.5 in) long. Like all tarsiers, the Philippine tarsier's eyes are fixed in its skull; they cannot move in their sockets. Instead, a special adaptation in the neck allows its round head to be rotated 180°. Their eyes are disproportionately large, having the largest eye-to-body weight ratio of all mammals. These huge eyes provide this nocturnal animal with excellent night vision. In bright light, the tarsier's eyes can constrict until the pupil appears to be only a thin spot. In low light or darkness, the pupil can dilate and fill up almost the entire eye. The large membranous ears are mobile, appearing to be almost constantly moving, allowing the tarsier to hear any movement.
The Philippine tarsier has thin, rough fur which is colored gray to dark brown. The narrow tail, usually used for balance, is bald except for a tuft of hair at the end, and is about twice the body length. Its elongated "tarsus", or ankle bone, which gives the tarsier its name, allows it to jump at least 3 m (9.8 ft) from tree to tree. Its long digits are tipped with rounded pads that allow C. syrichta to cling easily to trees and to grip almost any surface. The thumb is not truly opposable, but the first toe is. All of the digits have flattened nails, except for the second and third toes, which have sharp claws specialized for grooming. Their dental formula is 2:1:3:31:1:3:3, with relatively small upper canines.
Ecology
The Philippine tarsier is primarily insectivorous, its diet consists of insects, spiders, small crustaceans, and small vertebrates such as small lizards and birds. C. syrichta preys on live insects, particularly crickets and grasshoppers. Upon seizing its prey, the tarsier carries it to its mouth using both hands.
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