The piping plover (Charadrius melodus) is a small, sand-colored shorebird that nests and feeds along coastal sand and gravel beaches in North America. The adult has yellow-orange-red legs, a black band across the forehead from eye to eye, and a black stripe running along the breast line. This chest band is usually thicker in males during the breeding season, and it is the only reliable way to tell the sexes apart. The bird is difficult to see when it is standing still, as it blends well with open, sandy beach habitats. It typically runs in short, quick spurts and then stops. There are two subspecies of piping plovers: the eastern population is known as Charadrius melodus melodus and the mid-west population is known as C. m. circumcinctus. The bird's name is derived from its plaintive bell-like whistles which are often heard before the bird is visible. In 1986, the U.S. Fish & Wildlife Service listed the Great Lakes population as endangered and the Northern Great Plains and Atlantic populations as threatened. Intensive conservation efforts have yielded slow population growth, but only some populations have met the recovery goals set for them. The growth trend is expected to reverse if conservation efforts were stopped. Total population was estimated to be between 7600 and 8400 individuals in 2020. Their breeding habitat includes beaches and sand flats on the Atlantic coast, the shores of the Great Lakes, and in the mid-west of Canada and the United States. They nest on sandy or gravel beaches or shoals. These shorebirds forage for food on beaches, usually by sight, moving across the beaches in short bursts. Generally, piping plovers will forage for food around the high tide wrack zone and along the water's edge. They eat mainly insects, marine worms, and crustaceans.
Taxonomy
The piping plover was given the binomial name Charadrius melodus in 1824 by the American ornithologist George Ord in a revised edition of Alexander Wilson's American Ornithology; or, The Natural History of the Birds of the United States. In 1812 Wilson had described and illustrated a bird collected from Great Egg Harbour, New Jersey. Two subspecies are recognized: the nominate C. m. melodus of the Atlantic Coast and C. m. circumcinctus of the Great Plains. On average, circumcinctus is darker overall with more contrastingly dark cheeks and lores. Breeding circumcinctus males show more extensive black on forehead and bill-base and more often shows complete breast-bands. Some overlap exists.
Description
The piping plover is a stout bird with a large rounded head, a short thick neck, and a stubby bill. It is a sand-colored, dull gray/khaki, sparrow-sized shorebird. The adult has yellow-orange legs, the male has a prominent black band across the forehead from eye to eye, and a black ring around the neck during the breeding season. The band on the female's brow is much fainter. During nonbreeding season, the black bands become less pronounced. Its bill is orange with a black tip. It ranges from 15–19 cm (5.9–7.5 in) in length, with a wingspan of 35–41 cm (14–16 in) and a mass of 42–64 g (1.5–2.3 oz).
Vocalizations The piping plover's light call is a soft, whistled peep peep given by standing and flying birds. Its frequently heard alarm call is a soft pee-werp, which the second syllable lower pitched.
Distribution and habitat
The piping plover lives the majority of its life on open sandy beaches or rocky shores, often in high, dry sections away from water. They can be found on the Atlantic Coast of the U.S. and Canada on the ocean or bay beaches and on the Great Lakes shores. It builds its nests higher on the shore near beach grass and other objects. It is very rare to see a piping plover anywhere outside of sand or rocky beaches/shores while not migrating. Notably, they are considered a disturbance-dependent species due to their habitat's relationship with severe flooding. They need their nesting sites to be far away from the water to be dry for their entire nesting period – yet, they also need to nest in open sand. Often, this combination occurs only in sites which are thoroughly flooded once every few years, which brings in fresh sand and clears away vegetation. As human coastal management strategies worked to minimize unpredictable flooding, their old habitats often became overgrown, and their populations declined. Nowadays, the U.S. Army Corps of Engineers is forced to artificially clear shoals on the Platte and Missouri River to maintain some of the plovers' remaining habitat in the Great Plains. However, while these efforts are important, they remain inferior to natural periodically flooded habitats. Alongside least terns, piping plovers also have an unusual relationship with mining. Whereas most species predictably suffer habitat loss from mining activities in their area, these birds are known to nest in waste sand piles generated by nearby mines, as the replacement for sand bars. Those sand piles remain topped-up and hostile to vegetation overgrowth as long as the mining activity continues. Often, mining sites end up reclaimed for housing development at the end of their lifespan, which means more construction work, and an extended persistence of waste sand piles. However, modern mining practices typically transport much of this waste sand to more remote areas, often to reuse it commercially, which limits colonization opportunities for these birds. Piping plovers migrate from their northern range in the summer to the south in the winter months, migrating to the Gulf of Mexico, the southern Atlantic coast of the United States and the Caribbean, including The Bahamas. They have also been recorded across Cuba, with rarer occurrences elsewhere throughout the West Indies, and even Ecuador and Venezuela. They begin migrating north in mid-March. Their breeding grounds extend from southern Newfoundland south to the northern parts of South Carolina. Migration south begins in August for some adults and fledglings, and by mid-September most piping plovers have headed south for winter.
Behavior
Breeding
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