Although projectiles are commonly used in human conflict, projectile use by organisms other than humans is relatively rare. However, some organisms are capable of using various different types of projectiles for defense or predation. Some invertebrates are able to spit or squirt out fluids and some vertebrates also use liquid projectiles. Solid projectiles are used by various species. Tarantulas, for example, may kick off barbed hairs and several mammals have been observed to throw objects such as sand, rocks, sticks and feces. Chameleons, frogs and some salamanders have tongues that act like tethered projectiles. The pistol shrimp may emit a powerful wave of bubbles. Some plants produce seed pods that explode when touched or may fire spore capsules.
Animals
Liquid projectiles
Most projectiles used by terrestrial animals are liquids. Among invertebrates there are a number of examples. Velvet worms can squirt out a slimy adhesive fluid from glands on the sides of their head, and use it to trap their prey. The spitting spiders Scytodes can spit a venomous sticky fluid that traps its victims and also poisons them. The bombardier beetle is unusual by using a violent exothermic chemical reaction to launch a boiling noxious chemical spray in a rapid burst of pulses from special glands in its abdomen, accompanied with a popping sound. The Anthia (oogpister beetle) will fire formic acid at attackers, probably extracting the formic acid from the ants that it eats. The devil-rider stick insects (Anisomorpha) can fire terpenes from glands on the metathorax that can cause an intense burning irritation of the eyes and mouth of potential predators. Some species of ants of the subfamily Formicinae, like wood ants and weaver ants, will spray acid at attackers. They will bite to cause a wound and spray acid directly into the wound. A type of planthopper of Madagascar is able to flick small balls of honeydew, this attracts day geckos that feed on the honeydew and whose presence may deter predators from approaching the sap-sucking insect. Termites of the subfamily Nasutitermitinae can project a sticky fluid from a nozzle on their heads. They can use this fontanellar gun accurately, over a range of many centimeters, even though the termite is blind, possibly using auditory or olfactory cues instead. A number of vertebrates also use liquid projectiles. The archerfish will squirt water from its mouth to dislodge invertebrates from overhanging branches. Some diptodactyline geckos can fire a black or pale yellow sticky fluid out of glands in their tail for a distance of about a meter, and with good aim. This fluid has a musky unpleasant odour and although it is not toxic it may discourage predators, in particular the big arthropods that prey on these geckos. Some species of horned lizards (Phrynosoma) can fire an aimed stream of blood from the corners of the eyes for a distance up to 3 ft to deter predators. The spitting cobra can squirt venom from forward-facing holes in its fangs. It aims for its victim's eyes, spitting up to 1.5 meters (5 ft). The venom may cause blindness. The Mangshan pitviper is also reported to spit venom. A bird that uses liquid projectiles in defense is the southern giant petrel which produces a stomach oil made up of wax esters and triglycerides that are stored in the proventriculus and can be projectile vomited on predators. Other petrels such as the fulmar can also squirt oils from their mouths as a defense. They can squirt oils with accuracy up to a distance of 1 to 2 meters (3.5 to 6.5 ft). The oil mats the feathers of birds together and destroys their waterproofing abilities, so oiled birds may die from chilling or drowning, although fulmars seem able to remove the oil from themselves by preening. Birds ranging from gulls to sea-eagles have died after being squirted by fulmars. Some species of penguin expel liquid feces in a projectile manner, to a distance of up to about 50 cm (20 in). They are believed to do this because during the brooding season, when penguins are sitting on their nests, they avoid leaving their nests and thereby leaving their eggs open to predation and thus to maintain a clean nest they evolved the ability to project their feces. Among mammals, skunks can eject a noxious fluid from their anal glands. It is not only foul smelling, but can cause skin irritation and, if it gets in the eyes, temporary blindness. When it feels threatened a camel will bring up their stomach contents, along with saliva, and project it out towards the threat to distract, surprise, or bother the threat.
Solid projectiles
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