The quagga ( or ) (Equus quagga quagga) is an extinct subspecies of the plains zebra which was endemic to South Africa until it was hunted to extinction in the late 19th century. It was long thought to be a distinct species, but mtDNA studies have supported it being a subspecies of plains zebra. A more recent study suggested that it was the southernmost cline or ecotype of the species. The quagga is believed to have been around 257 cm (8 ft 5 in) long and 125–135 cm (4 ft 1 in – 4 ft 5 in) tall at the shoulders. It was distinguished from other zebras by its limited pattern of primarily brown and white stripes, mainly on the front part of the body. The rear was brown and without stripes, and appeared more horse-like. The distribution of stripes varied considerably between individuals. Little is known about the quagga's behaviour, but it may have gathered into herds of 30–50. Quaggas were said to be wild and lively, yet were also considered more docile than the related Burchell's zebra. They were once found in great numbers in the Karoo of Cape Province and the southern part of the Orange Free State in South Africa. After the European settlement of South Africa began, the quagga was extensively hunted, as it competed with domesticated animals for forage. Some were taken to zoos in Europe, but breeding programmes were unsuccessful. The last wild population lived in the Orange Free State; the quagga was extinct in the wild by 1878. The last captive specimen died in Amsterdam on 12 August 1883. 23 skins and seven skeletons exist today, and only two quaggas were ever photographed alive. In 1984, the quagga was the first extinct animal whose DNA was analysed. The Quagga Project has attempted to breed Burchell's zebras with similar striping patterns to the quagga.
Taxonomy It has been historically suggested that the name quagga is derived from the Khoekhoe word for zebra; cf. Kora !oaxais 'quagga', which is also the origin of the isiXhosa inqwarhashe 'quagga'.
The quagga was originally classified as a distinct species, Equus quagga, in 1778 by Dutch naturalist Pieter Boddaert. Traditionally, the quagga and the other plains and mountain zebras were placed in the subgenus Hippotigris. Much debate has occurred over the status of the quagga in relation to the plains zebra. The British zoologist Reginald Innes Pocock in 1902 was perhaps the first to suggest that the quagga was a subspecies of the plains zebra. As the quagga was scientifically described and named before the plains zebra, the trinomial name for the quagga becomes E. quagga quagga under this scheme, and the other subspecies of the plains zebra are placed under E. quagga, as well. Historically, quagga taxonomy was further complicated because the extinct southernmost population of Burchell's zebra (Equus quagga burchellii, formerly Equus burchellii burchellii) was thought to be a distinct subspecies (also sometimes thought a full species, E. burchellii). The extant northern population, the "Damara zebra", was later named Equus quagga antiquorum, which means that it is today also referred to as E. q. burchellii, after it was realised they were the same taxon. The extinct population was long thought very close to the quagga, since it also showed limited striping on its hind parts. As an example of this, Shortridge placed the two in the now disused subgenus Quagga in 1934. Most experts now suggest that the two subspecies represent two ends of a cline. Different subspecies of plains zebras were recognised as members of Equus quagga by early researchers, though much confusion existed over which species were valid. Quagga subspecies were described on the basis of differences in striping patterns, but these differences were since attributed to individual variation within the same populations. Some subspecies and even species, such as E. q. danielli and Hippotigris isabellinus, were based only on illustrations (iconotypes) of aberrant quagga specimens. One craniometric study from 1980 seemed to confirm its affiliation with the horse (Equus ferus caballus), but early morphological studies have been noted as being erroneous. Studying skeletons from stuffed specimens can be problematic, as early taxidermists sometimes used donkey and horse skulls inside their mounts when the originals were unavailable.
Evolution
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