The red wolf (Canis rufus) is a canine native to the southeastern United States. Its size is intermediate between the coyote (Canis latrans) and gray wolf (Canis lupus). The red wolf's taxonomic classification as being a separate species has been contentious for nearly a century, being considered either a subspecies of the gray wolf Canis lupus rufus, or a coywolf (a genetic admixture of wolf and coyote). Because of this, it is sometimes excluded from endangered species lists, despite its critically low numbers. Under the Endangered Species Act of 1973, the U.S. Fish and Wildlife Service recognizes the red wolf as an endangered species and grants it protected status. Since 1996, the IUCN has listed the red wolf as a Critically Endangered species; however, it is not listed in the CITES Appendices of endangered species.
History Red wolves were once distributed throughout the southeastern and south-central United States from the Atlantic Ocean to central Texas, southeastern Oklahoma and southwestern Illinois in the west, and in the north from the Ohio River Valley, northern Pennsylvania, southern New York, and extreme southern Ontario in Canada south to the Gulf of Mexico. The red wolf was nearly driven to extinction by the mid-1900s due to aggressive predator-control programs, habitat destruction, and extensive hybridization with coyotes. By the late 1960s, it occurred in small numbers in the Gulf Coast of western Louisiana and eastern Texas. Fourteen of these survivors were selected to be the founders of a captive-bred population, which was established in the Point Defiance Zoo and Aquarium between 1974 and 1980. After a successful experimental relocation to Bulls Island off the coast of South Carolina in 1978, the red wolf was declared extinct in the wild in 1980 so that restoration efforts could proceed. In 1987, the captive animals were released into the Alligator River National Wildlife Refuge (ARNWR) on the Albemarle Peninsula in North Carolina, with a second unsuccessful release taking place two years later in the Great Smoky Mountains National Park. Of 63 red wolves released from 1987 to 1994, the population rose to as many as 100–120 individuals in 2012, but due to the lack of regulation enforcement by the US Fish and Wildlife Service, the population has declined to 40 individuals in 2018, about 14 in 2019 and 8 as of October 2021. No wild litters were born between 2019 and 2020. Under pressure from conservation groups, the US Fish and Wildlife Service resumed reintroductions in 2021 and increased protection. In 2022, the first wild litter was born since 2018. As of 2023, there are between 15 and 17 wild red wolves in ARNWR.
Description and behavior
The red wolf's appearance is typical of the genus Canis, and is generally intermediate in size between the coyote and gray wolf, though some specimens may overlap in size with small gray wolves. A study of Canis morphometrics conducted in eastern North Carolina reported that red wolves are morphometrically distinct from coyotes and hybrids. Adults measure 136–165 cm (53.5–65 in) in length, comprising a tail of about 37 cm (14.6 in). Their weight ranges from 20 to 39 kg (44–85 lbs) with males averaging 29 kg (64 lbs) and females 25 kg (55 lbs). Its pelage is typically more reddish and sparsely furred than the coyote's and gray wolf's, though melanistic individuals do occur. Its fur is generally tawny to grayish in color, with light markings around the lips and eyes. The red wolf has been compared by some authors to the greyhound in general form, owing to its relatively long and slender limbs. The ears are also proportionately larger than the coyote's and gray wolf's. The skull is typically narrow, with a long and slender rostrum, a small braincase and a well developed sagittal crest. Its cerebellum is unlike that of other Canis species, being closer in form to that of canids of the Vulpes and Urocyon genera, thus indicating that the red wolf is one of the more plesiomorphic members of its genus. The red wolf is more sociable than the coyote, but less so than the gray wolf. It mates in January–February, with an average of 6–7 pups being born in March, April, and May. It is monogamous, with both parents participating in the rearing of young. Denning sites include hollow tree trunks, along stream banks and the abandoned earths of other animals. By the age of six weeks, the pups distance themselves from the den, and reach full size at the age of one year, becoming sexually mature two years later. Using long-term data on red wolf individuals of known pedigree, it was found that inbreeding among first-degree relatives was rare. A likely mechanism for avoidance of inbreeding is independent dispersal trajectories from the natal pack. Many of the young wolves spend time alone or in small non-breeding packs composed of unrelated individuals. The union of two unrelated individuals in a new home range is the predominant pattern of breeding pair formation. Inbreeding is avoided because it results in progeny with reduced fitness (inbreeding depression) that is predominantly caused by the homozygous expression of recessive deleterious alleles. Prior to its extinction in the wild, the red wolf's diet consisted of rabbits, rodents, and nutria (an introduced species). In contrast, the red wolves from the restored population rely on white-tailed deer, pig, raccoon, rice rats, muskrats, nutria, rabbits and carrion. White-tailed deer were largely absent from the last wild refuge of red wolves on the Gulf Coast between Texas and Louisiana (where specimens were trapped from the last wild population for captive breeding), which likely accounts for the discrepancy in their dietary habits listed here. Historical accounts of wolves in the southeast by early explorers such as William Hilton, who sailed along the Cape Fear River in what is now North Carolina in 1644, also note that they ate deer.
Range and habitat
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