Tanystropheus (Ancient Greek: τανυ~ 'long' + στροφευς 'hinged') is an extinct genus of archosauromorph reptile which lived during the Triassic Period in Europe, Asia, and North America. It is recognizable by its extremely elongated neck, longer than the torso and tail combined. The neck is composed of 13 vertebrae, strengthened by long cervical ribs. Tanystropheus is one of the most well-described non-archosauriform archosauromorphs (reptiles closer to dinosaurs and crocodilians than to lizards). It is known from numerous fossils, including nearly complete skeletons. Some species within the genus may have reached a total length of 6 meters (20 ft), making Tanystropheus the longest non-archosauriform archosauromorph as well. Tanystropheus is the namesake of the family Tanystropheidae, a clade collecting many long-necked Triassic archosauromorphs previously described as "protorosaurs" or "prolacertiforms". Tanystropheus contains at least two valid species as well as fossils which cannot be referred to a specific species. The type species of Tanystropheus is T. conspicuus, a dubious name applied to particularly large fossils from Germany and Poland. Complete skeletons are common in the Besano Formation at Monte San Giorgio, on the border of Italy and Switzerland. Monte San Giorgio fossils belong to two species: the smaller T. longobardicus and the larger T. hydroides. When T. longobardicus was first described in 1886, it was initially mistaken for a pterosaur and given the name "Tribelesodon". Starting in the 1920s, systematic excavations at Monte San Giorgio unearthed many more Tanystropheus fossils, revealing that the supposed wing bones of "Tribelesodon" were actually neck vertebrae. T. hydroides was formally distinguished in 2020, mostly on the basis of its distinctive skull and the confirmation that the two different size forms are adults of different species. Most Tanystropheus fossils hail from marine or coastal deposits of the Middle Triassic epoch (Anisian and Ladinian stages), with some exceptions. For example, a vertebra from Nova Scotia was recovered from primarily freshwater sediments. The youngest unambiguous fossils in the genus are a pair of well-preserved skeletons from the Zhuganpo Formation, a geological unit in China which dates to the earliest part of the Late Triassic (early Carnian stage). The oldest claims of Tanystropheus fossils belong to "T. antiquus", a European species from the latest part of the Early Triassic (late Olenekian stage). T. antiquus has a proportionally shorter neck than other Tanystropheus species, so some paleontologists consider that T. antiquus deserves a separate genus, Protanystropheus. Some studies suggest that T. antiquus is not even part of the family Tanystropheidae, instead being closer to another long-necked early archosauromorph, Dinocephalosaurus. The lifestyle of Tanystropheus has been the subject of much debate. Tanystropheus is unknown from drier environments and its neck is rather stiff and ungainly, suggesting a reliance on water. On the other hand, the limbs and tail lack most adaptations for swimming and are similar to those of land-based reptiles. Recent studies have supported an intermediate position, reconstructing Tanystropheus as an animal equally capable on land and in the water. Despite its length, the neck was lightweight and stabilized by tendons, so it would not have been a fatal hindrance to moving on land. The hindlimbs and the base of the tail were large and muscular, capable of short bursts of active swimming in shallow water. Tanystropheus was most likely a piscivorous (fish-eating) ambush predator. The narrow and tapering skull of T. longobardicus has three-cusped teeth at the back, suited for holding onto slippery prey, while the broader skull of T. hydroides has an interlocking set of large curved fangs at the front, similar to the fully aquatic plesiosaurs.
History and species
Monte San Giorgio species
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