This timeline of mosasaur research is a chronologically ordered list of important fossil discoveries, controversies of interpretation, and taxonomic revisions of mosasaurs, a group of giant marine lizards that lived during the Late Cretaceous Epoch. Although mosasaurs went extinct millions of years before humans evolved, humans have coexisted with mosasaur fossils for millennia. Before the development of paleontology as a formal science, these remains would have been interpreted through a mythological lens. Myths about warfare between serpentine water monsters and aerial thunderbirds told by the Native Americans of the modern western United States may have been influenced by observations of mosasaur fossils and their co-occurrence with creatures like Pteranodon and Hesperornis. The scientific study of mosasaurs began in the late 18th century with the serendipitous discovery of a large fossilized skeleton in a limestone mine near Maastricht in the Netherlands. The fossils were studied by local scholar Adriaan Gilles Camper, who noted a resemblance to modern monitor lizards in correspondence with renowned French anatomist Georges Cuvier. Nevertheless, the animal was not scientifically described until the English Reverend William Daniel Conybeare named it Mosasaurus, after the river Meuse located near the site of its discovery. By this time the first mosasaur fossils from the United States were discovered by the Lewis and Clark Expedition, and the first remains in the country to be scientifically described were reported slightly later from New Jersey. This was followed by an avalanche of discoveries by the feuding Bone War paleontologists Edward Drinker Cope and Othniel Charles Marsh in the Smoky Hill Chalk of Kansas. By the end of the century a specimen of Tylosaurus would be found that preserved its scaley skin. Later Samuel Wendell Williston mistook fossilized tracheal rings for the remains of a fringe of skin running down the animal's back, which subsequently became a common inaccuracy in artistic restorations. The 20th century soon saw the discovery in Alabama of a strange mosasaur called Globidens, with rounded teeth suited to crushing shells. Mosasaur remains were also discovered in Africa and California. In 1967 Dale Russell published a scientific monograph dedicated to mosasaurs. Embryonic remains in the 1990s confirmed that mosasaurs gave live birth like in ichthyosaurs. The 1990s also saw a revival and escalation of a debate regarding whether or not some supposed mosasaur toothmarks in ammonoid shells were actually made by limpets. By the end of the century, the evolutionary relationship between mosasaurs and snakes as well as the possible involvement of mosasaurs in the extinction of the aforementioned ichthyosaurs became hot button controversies. The debates regarding snakes, toothmarks, and ichthyosaurs spilled over into the early 21st century. These discussions were also accompanied by the discovery of many new taxa, including new species of Globidens, Mosasaurus, and Tylosaurus as well as entirely new genera like Yaguarasaurus and Tethysaurus. In 2013, Lindgren, Kaddumi, and Polcyn reported the discovery of a Prognathodon specimen from Jordan that preserved the soft tissues of its scaley skin, flippers and tail. Significantly, the tail resembled those of modern carcharinid sharks, although the bottom lobe of the tail fin was longest in the mosasaur whereas shark tails have longer upper lobes.
Prescientific
One legend told among the Lenape (Delaware) people describes the discovery of a fragment of bone left by a monster that had once been killing people near modern Philadelphia in either Pennsylvania or eastern New Jersey. When the hunting party brought the piece of bone back to the village, a wise man encouraged them to set out and find more of the monsters remains. He said that smoking fragments of the bone with tobacco in a small clay spoon could grant wishes like good health for one's children, longevity, or successful hunting. Fossils of the mosasaur Tylosaurus have been found in the area, but dinosaurs and Cretaceous crocodiles are also possible sources. This legend likely predates European contact and may have originated prior to 1500. The bones being burnt in the "clay spoon" is a reference to a primitive kind of clay pipe that the local people had abandoned in favor of a design with a deeper bowl by the 17th century. The general lack of any sign of influence by European culture on the tale is also suggestive of its great antiquity. The Cheyenne people of Nebraska believed in mythical thunderbirds and water monsters that were in endless conflict with each other. The thunderbirds were said to resemble giant eagles and killed both people and animals with arrows made of lightning. The fossils of the Niobrara Chalk may have been influential on these stories. The pterosaur Pteranodon and marine reptiles like mosasaurs are preserved in Niobrara Chalk deposits and associated remains may have been interpreted as evidence for antagonism between immense flying animals and serpentine aquatic reptiles. Observations of similar fossils in the past may have been seen as further evidence for thunderbird-water monster conflict. The Cheyenne believe that there were many different kinds of water monsters that lived not only in lakes, rivers, and springs but also high bluffs and hills. The locations given as water monster habitat are similar to the locations where local marine fossils can be found as fossils often erode out of hillsides or stream banks. The Cheyennes feared the water monsters, because they could be dangerous predators or capsize their canoes. Even in modern times, tradition-minded Cheyenne sometimes take pains to avoid sleeping too close to springs due to fears of water monsters.
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