The tongue is a muscular organ in the mouth of a typical tetrapod. It manipulates food for chewing and swallowing as part of the digestive process and is the primary organ of taste. The tongue's upper surface (dorsum) is covered by taste buds housed in numerous lingual papillae. It is sensitive and kept moist by saliva and is richly supplied with nerves and blood vessels. The tongue also serves as a natural means of cleaning the teeth. A major function of the tongue is to enable speech in humans and vocalization in other animals. The human tongue is divided into two parts, an oral part at the front and a pharyngeal part at the back. The left and right sides are also separated along most of its length by a vertical section of fibrous tissue (the lingual septum) that results in a groove, the median sulcus, on the tongue's surface. There are two groups of glossal muscles. The four intrinsic muscles alter the shape of the tongue and are not attached to bone. The four paired extrinsic muscles change the position of the tongue and are anchored to bone.
Etymology The word tongue derives from the Old English tunge, which comes from Proto-Germanic *tungōn. It has cognates in other Germanic languages—for example tonge in West Frisian, tong in Dutch and Afrikaans, Zunge in German, tunge in Danish and Norwegian, and tunga in Icelandic, Faroese and Swedish. The ue ending of the word seems to be a fourteenth-century attempt to show "proper pronunciation", but it is "neither etymological nor phonetic". Some used the spelling tunge and tonge as late as the sixteenth century.
In humans
Structure
The tongue is a muscular hydrostat that forms part of the floor of the oral cavity. The left and right sides of the tongue are separated by a vertical section of fibrous tissue known as the lingual septum. This division is along the length of the tongue save for the very back of the pharyngeal part and is visible as a groove called the median sulcus. The human tongue is divided into anterior and posterior parts by the terminal sulcus, which is a "V"-shaped groove. The apex of the terminal sulcus is marked by a blind foramen, the foramen cecum, which is a remnant of the median thyroid diverticulum in early embryonic development. The anterior oral part is the visible part situated at the front and makes up roughly two-thirds the length of the tongue. The posterior pharyngeal part is the part closest to the throat, roughly one-third of its length. These parts differ in terms of their embryological development and nerve supply. The anterior tongue is, at its apex, thin and narrow. It is directed forward against the lingual surfaces of the lower incisor teeth. The posterior part is, at its root, directed backward, and connected with the hyoid bone by the hyoglossi and genioglossi muscles and the hyoglossal membrane, with the epiglottis by three glossoepiglottic folds of mucous membrane, with the soft palate by the glossopalatine arches, and with the pharynx by the superior pharyngeal constrictor muscle and the mucous membrane. It also forms the anterior wall of the oropharynx. The average length of the human tongue from the oropharynx to the tip is 10 cm. The average weight of the human tongue from adult males is 99g and for adult females 79g. In phonetics and phonology, a distinction is made between the tip of the tongue and the blade (the portion just behind the tip). Sounds made with the tongue tip are said to be apical, while those made with the tongue blade are said to be laminal. Tongue posture is the resting position of the tongue in the mouth. Evidence demonstrates that the tongue plays a role in mouth and face development.
Upper surface
The upper surface of the tongue, the dorsal surface, is called the dorsum, and is divided by a groove into symmetrical halves by the median sulcus. The foramen cecum marks the end of this division (at about 2.5 cm from the root of the tongue) and the beginning of the terminal sulcus. The foramen cecum is also the point of attachment of the thyroglossal duct and is formed during the descent of the thyroid diverticulum in embryonic development. The terminal sulcus is a shallow groove that runs forward as a shallow groove in a V shape from the foramen cecum, forwards and outwards to the margins (borders) of the tongue. The terminal sulcus divides the tongue into a posterior pharyngeal part and an anterior oral part. The pharyngeal part is supplied by the glossopharyngeal nerve and the oral part is supplied by the lingual nerve (a branch of the mandibular branch (V3) of the trigeminal nerve) for somatosensory perception and by the chorda tympani (a branch of the facial nerve) for taste perception. Both parts of the tongue develop from different pharyngeal arches.
Undersurface On the undersurface, the ventral surface, of the tongue is a fold of mucous membrane called the frenulum that tethers the tongue at the midline to the floor of the mouth. On either side of the frenulum are small prominences called sublingual caruncles that the major salivary submandibular glands drain into.
Muscles The eight muscles of the human tongue are classified as either intrinsic or extrinsic. The four intrinsic muscles act to change the shape of the tongue, and are not attached to any bone. The four extrinsic muscles act to change the position of the tongue, and are anchored to bone.
Extrinsic
The four extrinsic muscles originate from bone and extend to the tongue. They are the genioglossus, the hyoglossus (often including the chondroglossus) the styloglossus, and the palatoglossus. Their main functions are altering the tongue's position allowing for protrusion, retraction, and side-to-side movement. The genioglossus arises from the mandible and protrudes the tongue. It is also known as the tongue's "safety muscle" since it is the only muscle that propels the tongue forward. The hyoglossus, arises from the hyoid bone and retracts and depresses the tongue. The chondroglossus is often included with this muscle. The styloglossus arises from the styloid process of the temporal bone and draws the sides of the tongue up to create a trough for swallowing. The palatoglossus arises from the palatine aponeurosis, and depresses the soft palate, moves the palatoglossal fold towards the midline, and elevates the back of the tongue during swallowing.
Intrinsic
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