Tylopterella is a genus of eurypterid, a group of extinct aquatic arthropods. Only one fossil of the single and type species, T. boylei, has been discovered in deposits of the Late Silurian period (Ludlow epoch) in Elora, Canada. The name of the genus is composed by the Ancient Greek words τύλη (túlē), meaning "knot", and πτερόν (pteron), meaning "wing". The species name boylei honors David Boyle, who discovered the specimen of Tylopterella. It is a poorly-known genus whose carapace (dorsal plate of the prosoma, head) was semiovate bordered by a marginal rim, with eyes laterally placed, a preabdomen and postabdomen (the two halves of the abdomen) with six segments each and a short spike-like telson (which was the posteriormost division of the body). It reached a total length of 7.5 centimetres (2.9 inches). These characteristics place Tylopterella in the family Onychopterellidae together with Onychopterella. Tylopterella is notable for its thick ornamentation and general body surface. Its paired tubercles or knobs in the top of its second to fifth segments differentiates it from many other eurypterids. This thickness that its body possessed is due to the highly saline conditions to which Tylopterella had to adapt in the Guelph Formation; other organisms with reinforced shells have also been found in the same place.
Description
Like the other onychopterellids, Tylopterella was a small eurypterid. The total size of the only known specimen is estimated at only 7.5 centimetres (2.9 inches). Tylopterella is a little-known genus; the type specimen of T. boylei, which conserves the dorsal part of the body, represents its only record thus far. The carapace (dorsal plate of the prosoma, head) had a semiovate (nearly oval) shape, was rounded anteriorly and truncated (shortened as by cutting it) posteriorly and bordered by a highly raised, narrow rim more marked at the sides of the carapace. It was wider than long, and was notable for the extraordinary thickness of its surface and ornamentation. The carapace was 2 cm long (0.8 in) and 2.7 cm wide (1.1 in). The eyes were more or less laterally placed, reniform (bean-shaped) and prominent, with about 0.4 cm (0.16 in) in the greatest diameter and separated between them by 0.6 cm (0.23 in). Equidistantly from the eyes (that is, as far from one eye as from the other), there was a small rounded prominence in which the ocelli (simple eye-like sensory organs) were located. The surface of the carapace was granulate (with granules) and had an ornamentation which consisted of minute rounded tubercles, some isolated and others confluent in sets of two or three. It was strengthened by prominent calcareous (with calcium) deposits. As in the rest of eurypterids, the opisthosoma (abdomen) had twelve segments. It was divided into the preabdomen (segments 1 to 6), which was telescoped (with segments overlapping each other) and postabdomen (segments 7 to 12). It was thick, short and compact, specially in the two last segments. It was covered by a calcareo-chitinous (with chitin) layer. Each of the second to fifth tergites (dorsal half of the segment) carried in its median line a pair of single large, solid, prominent and elongated tubercles or knobs. They were reniform at the base and somewhat bilobed (with two lobes) at the top. The telson (the posteriormost division of the body) was like a gradually narrowing, slightly curved and pointed spine. It reached a length of 1.5 cm (0.6 in). Fragments of the fourth appendage (limb) and the paddle of the swimming leg (sixth and last pair of appendages) are also preserved.
History of research
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