Wheat is a group of wild and domesticated grasses of the genus Triticum (). As cereals, they are cultivated for their grains, which are staple foods around the world. Well-known wheat species and hybrids include the most widely grown common wheat (T. aestivum), spelt, durum, emmer, einkorn, and Khorasan or Kamut. The archaeological record suggests that wheat was first cultivated in the regions of the Fertile Crescent around 9600 BC. Wheat is grown on a larger area of land than any other food crop (219.5 million hectares or 542 million acres in 2024). World trade in wheat is greater than that of all other crops combined. In 2024, world wheat production was 799 million tonnes (881 million short tons), making it the second most-produced cereal after maize (known as corn in North America and Australia). Since 1960, world production of wheat and other grain crops has tripled and is expected to grow further through the middle of the 21st century. Global demand for wheat is increasing because of the usefulness of gluten to the food industry. Wheat is an important source of carbohydrates. Globally, it is the leading source of vegetable proteins in human food, having a protein content of about 13%, which is relatively high compared to other major cereals but relatively low in protein quality (supplying essential amino acids). When eaten as the whole grain, wheat is a source of multiple nutrients and dietary fibre. In a small part of the general population, gluten – which comprises most of the protein in wheat – can trigger coeliac disease, noncoeliac gluten sensitivity, gluten ataxia, and dermatitis herpetiformis.
Description
Wheat is a stout grass of medium to tall height. Its stem is jointed and usually hollow, forming a straw. There can be many stems on one plant. It has long narrow leaves, their bases sheathing the stem, one above each joint. At the top of the stem is the flower head, containing some 20 to 100 flowers. Each flower contains both male and female parts. The flowers are wind-pollinated, with over 99% of pollination events being self-pollinations and the rest cross-pollinations. The flower is housed in a pair of small leaflike glumes. The two (male) stamens and (female) stigmas protrude outside the glumes. The flowers are grouped into spikelets, each with between two and six flowers. Each fertilised carpel develops into a wheat grain or berry; botanically a caryopsis fruit, it is often called a seed. The grains ripen to a golden yellow; a head of grain is called an ear. Leaves emerge from the shoot apical meristem in a telescoping fashion until the transition to reproduction i.e. flowering. The last leaf produced by a wheat plant is known as the flag leaf. It is denser and has a higher photosynthetic rate than other leaves, to supply carbohydrate to the developing ear. In temperate countries the flag leaf, along with the second and third highest leaves on the plant, supply the majority of carbohydrate in the grain; their condition is critical for crop yield. Wheat is unusual in having more stomata on the upper (adaxial) side of the leaf, than on the under (abaxial) side. It has been theorised that this might be an effect of having been cultivated longer than any other plant. Winter wheat generally produces up to 15 leaves per shoot, and spring wheat up to 9; winter crops may have up to 35 tillers (shoots) per plant (depending on cultivar). Wheat roots are among the deepest of arable crops, extending as far down as 2 metres (6 ft 7 in). While the roots of a wheat plant are growing, the plant accumulates an energy store in its stem, in the form of fructans, which helps the plant to yield under drought and disease pressure, but there is a trade-off between root growth and stem non-structural carbohydrate reserves. Root growth is likely to be prioritised in drought-adapted crops, while stem non-structural carbohydrate is prioritised in varieties developed for countries where disease is a bigger issue. Depending on variety, wheat may be awned or not. Producing awns incurs a cost in grain number, but wheat awns photosynthesise more efficiently than leaves with regards to water usage, so awns are much more frequent in varieties of wheat grown in hot drought-prone countries than those in temperate countries. For this reason, awned varieties could become more widespread due to climate change. In Europe, wheat's climate resilience has declined.
History
Domestication
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