The Wheeler Shale (named by Charles Walcott) is a Cambrian (c. 507 Ma) fossil locality world-famous for prolific agnostid and Elrathia kingii trilobite remains (even though many areas are barren of fossils) and represents a Konzentrat-Lagerstätte. Varied soft bodied organisms are locally preserved, a fauna (including Naraoia, Wiwaxia and Hallucigenia) and preservation style (carbonaceous film) normally associated with the more famous Burgess Shale. As such, the Wheeler Shale also represents a Konservat-Lagerstätten. Together with the Marjum Formation and lower Weeks Formation, the Wheeler Shale forms 490 to 610 m (1,610 to 2,000 ft) of limestone and shale exposed in one of the thickest, most fossiliferous and best exposed sequences of Middle Cambrian rocks in North America. At the type locality of Wheeler Amphitheater, House Range, Millard County, western Utah, the Wheeler Shale consists of a heterogeneous succession of highly calcareous shale, shaley limestone, mudstone and thin, flaggy limestone. The Wheeler Formation (although the Marjum & Weeks Formations are missing) extends into the Drum Mountains, northwest of the House Range where similar fossils and preservation are found.
Taphonomy and sedimentology
Detailed work recognises a number of ~10 m thick lagerstätten sequences in the formation, each of which formed at a sea-level highstand in deep water. The lagerstätte were deposited by turbidities and mudslides onto an oxygenated sea floor. The productive layers comprise mud and clay particles, with a tiny fraction of wind-blown quartz.
Stratigraphy The Wheeler Shale spans the Ptychagnostus atavus and uppermost-Middle Cambrian Bolaspidella trilobite zones (See House Range for full stratigraphy).
Fauna
Incomplete list of the fauna of the Wheeler Shale: (Note: the preservation of hard bodied trilobite remains and soft bodied animals seems to be mutually exclusive within particular horizons.)
Protista Marpolia spissa - cyanobacteria or green algae Morania fragmenta - cyanobacteria
Arthropoda Alalcomenaeus cambricus - megacheiran Branchiocaris pretiosa - hymenocarine Branchiocaris sp. Cambropodus gracilis - possible myriapod Canadaspis perfecta - hymenocarine Dicerocaris opisthoeces Dicranocaris guntherorum - possible habeliid? Emeraldella brocki - vicissicaudatan Isoxys Leanchoilia superlata - megacheiran Megachelicerax - chelicerate Mollisonia symmetrica - mollisoniid chelicerate Perspicaris dilatus hymenocarine Pseudoarctolepis sharpi - possible hymenocarine Sidneyia inexpectans - vicissicaudatan Tuzoia? peterseni - hymenocarine Waptia fieldensis - hymenocarine
Dinocaridida Amplectobelua cf. A. stephenensis – radiodont Anomalocarididae gen. et sp. nov. - radiodont Buccaspinea cooperi? - radiodont Caryosyntrips durus - radiodont Caryosyntrips serratus - radiodont Peytoia nathorsti - radiodont Pahvantia hastata - radiodont Stanleycaris sp. - radiodont Utahnax vannieri - kerygmachelid lobopodian, possibly from Marjum Formation Utaurora comosa - opabiniid
Trilobita Naraoia compacta - naraoiid nectaspid Hypagnostus parvifrons - agnostid Peronopsis amplaxis - peronopsid agnostid Peronopsis bidens Peronopsis fallax Peronopsis gaspensis Peronopsis intermedius Peronopsis interstrictus Peronopsis montis Peronopsis segmentis Ptychagnostus atavus (= Acidusus atavus) - ptychagnostid agnostid Ptychagnostus germanus Ptychagnostus gibbus Ptychagnostus intermedius Ptychagnostus michaeli Ptychagnostus occultatus Ptychagnostus seminula Glyphaspis concavus - asaphid Bathyuriscus fimbriatus - dolichometopid corynexochid Bathyuriscus sp. Kootenia sp. - dorypygid corynexochid, perhaps a synonym of Olenoides Olenoides expansus - dorypygid corynexochid Olenoides nevadensis Olenoides serratus Tonkinella breviceps Zacanthoides divergens - zacanthoidid corynexochid Zacanthoides sp. Altiocculus harrisi - ptychopariid (specific name may be confused with Alokistocare) Alokistocare harrisi - alokistocarid ptychopariid Asaphiscus wheeleri - ptychopariid; second-most common species in the formation Bathyocos housensis - ptychopariid Bolaspidella drumensis Bolaspidella housensis Bolaspidella sp. Bolaspidella wellsvillensis Brachyaspidion microps Brachyaspidion sulcatum Cedaria minor - known from the Warrior Formation Elrathia kingii - alokistocarid ptychopariid Elrathia sp. Elrathina wheeleri = Ptychoparella wheeleri? - ptychopariid Jenkinsonia varga Modocia brevispina Modocia laevinucha Modocia typicalis Ptychoparella sp. - ptychopariid Ptychoparella wheeleri Spencella sp. - ptychopariid
Brachiopoda Acrothele subsidua
Chordata Hertzina sp. - conodont
Cnidaria Cambromedusa sp. - jellyfish
Mollusca Pelagiella sp. - pelagiellid helcionelloid
Echinodermata Castericystis sprinklei - solute Castericystis sp. Cothurnocystis sp. - stylophoran Ctenocystis sp. - ctenocystoid Gogia spiralis - eocrinoid Eocrinoid holdfasts believed to belong to Gogia spiralis; may belong to other species
Porifera Choia carteri - choiid protomonaxonid demosponge Choia utahensis Crumillospongia sp. - hazeliid protomonaxonid demosponge Diagonella sp.
Priapulida Ottoia prolifica - archaeopriapulid Selkirkia sp. - archaeopriapulid Selkirkia willoughbyi- archaeopriapulid
Unclassified Hallucigenia sparsa - hallucigeniid lobopod Allonnia cf. tintinopsis - a chancelloriid Chancelloria pentacta - chancelloriid Eldonia sp. - eldoniid cambroernid Skeemella clavula - Possible vetulicolian Hylolithellus sp. - annelid? Wiwaxia corrugata - halwaxiid? lophotrochozoan Yuknessia simplex - pterobranch Margaretia dorus - tube associated with the hemichordate Oesia
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